King et al. 2019


Jolles et al. 2020

Tuci et al. 2019

Walker et al. 2017

King & Cowlishaw 2009

Sueur et al. 2011

King et al. 2009

King & Sueur 2011

Current Biology

King et al. 2008

King & Cowlishaw 2009

This is a publication list for PI Andrew, with SHOAL authors in bold textFor individual publication lists, follow links provided in our members page. All accessible [data and code] or [media coverage and videos] are also provided with publications.



[71] Fürtbauer, I., Christensen, C., Bracken, A., O'Riain, M.J., Heistermann, M., King, A.J. (in press) Energetics at the urban edge: Environmental and individual predictors of urinary C-peptide levels in wild chacma baboons (Papio ursinus). Hormones and Behavior

[70]  Fehlmann, G., O’Riain, M. J., Fürtbauer, I., King, A. J. (in press) Behavioural causes, ecological consequences, and management challenges associated with wildlife foraging in human-modified landscapes. BioScience

[69] Rafiq, K., Pitcher, B. J., Cornelsen, K., Hansen, W. K., King, A. J., Appleby, R. G., Abrahms, B., Jordan, N. R. (in press) Animal-borne technologies in wildlife research and conservation. In: Conservation Technology, Piel, A., Wich, S (Eds).

[68] Marsden, K. A., Lush, L., Holmberg, J. A., Charteris, A., Cardenas, L., King, A. J., Wilson, R. P., Jones, D. L., Chadwick, D. R. (2020) Sheep urination frequency, volume, N excretion and chemical composition: implications for subsequent agricultural N losses. Agriculture, Ecosystems & Environment online early 

[67] Zhang, J. King, A. J., Fürtbauer, I., Wang, Y-W., He, Y-Q., Zhang, Z-W., Wan, D-M., Yin, Y-X. (2020) Facilitative effects of social partners on Java sparrow activity. Animal Behaviour 161: 33-38



[66] Jolles, J. W., King, A. J., Killen, S. (2019) The role of individual heterogeneity in collective animal behaviour. Trends in Ecology & Evolution 35: 278-291


[65] Tuci, E., Trianni, V., King, A. J., Garnier, S. (2019) Editorial: Novel technological and methodological tools for the understanding of collective behaviours. Frontiers in Robotics and AI  6:139 


[64] de la O, C., Fürtbauer, I., King, A. J., Valenzuela-Galván, D. (2019) A resident-nepotistic-tolerant dominance style in wild white-nosed coatis (Nasua narica)? Behaviour 156(9)

[63] O'Bryan, L. R., Abaid, N., Nakayama, S., Dey, T., King, A. J., Cowlishaw, G., Rubenstein, D., Garnier, S. (2019) Contact calls facilitate group contraction in free-ranging goats (Capra aegagrus hircus). Frontiers Ecology and Evolution 7(Article 73) [data]



[62] Williams, H. J., King, A. J., Duriez, O., Börger, L., Shepard, E. L. C. (2018) Social eavesdropping allows for a more risky gliding strategy by thermal-soaring birds. Journal of the Royal Society Interface  15: 20180578 [Data - Figshare] [Data - Movebank 'Gyps Vultures']

[61] Ramos-Fernandez, G., King, A. J., Beehner, J. C., Bergman, T. J., Crofoot, M. C., Di Fiore, A., Lehmann, J. Schafner, C. M., Snyder-Mackler, N., Zuberbühler, K., Aureli, F., Boyer, D. (2018) Quantifying uncertainty due to fission-fusion dynamics as a component of social complexity. Proceedings of the Royal Society B: Biological Sciences 285: 1879[Data - Dryad]   

[60] Lush, L., Chadwick, D.,  Hopkins, P., Marsden, K., Holton, M., Wilson, R. P., King, A. J. (2018) Classification of sheep urination events using accelerometers to aid improved measurements of livestock contributions to nitrous oxide emissions.  Computers and Electronics in Agriculture 150: 170–177. 

[59] Strömbom, D. , King, A. J. (2018) Robot collection and transport of objects: a biomimetic process. Frontiers in Robotics and AI, section Evolutionary Robotics 5: 48 [The Conversation] [Video: robot trials]

[58] King, A. J., Fehlmann, G., Biro, D. Ward, A.J., Fürtbauer, I. (2018) Re-wilding collective behaviour: an ecological perspective.  Trends in Ecology & Evolution 33: 349-359 [Cover image] [Electronics360] [Science News Online]


[57] Fehlmann, G., O’Riain, M. J., Kerr-Smith, C., Hailes, S., Luckman, A., Shepard, E. L. C., King, A. J. (2017) Extreme behavioural shifts by baboons exploiting risky, resource-rich, human-modified environments. Scientific Reports 7:15057 [Supplementary material & R code] [Data - Dryad] [Video: summary] [TimesLive - South Africa] [Newsweek - US] [Der Standard - Austria] [Telegraph - India]

[56] Walker, R. H., King A. J., McNutt, J. W., Jordan, N. R. (2017) Sneeze to leave:  African wild dogs (Lycaon pictus) use variable quorum thresholds facilitated by sneezes in collective decisions Proceedings of the Royal Society B: Biological Sciences  284: 20170347  [Data - Dryad]. [Cover Image] Some(!) of the news: [ITV News] [BBC News] [Earth Times] [Seeker] [Cosmos] [New Atlas] [International Business Times] [National Geographic] [The Times] [New York Times] [Freakanomics

[55] King, A. J., Kosfeld, M., Dall, S. R. X., Greiner, B., Kameda, T., Khalmetski, K., Leininger, W., Wedekind, C., & Winterhalder, B. (2017). Exploitative Strategies: Consequences for Individual Behavior, Social Structure, and Design of Institutions. In: Investors and Exploiters in Ecology and Economics: Principles and Applications. Eds. L-A. Giraldeau, P. Heeb, M. Kosfeld. MIT Press.

[54] King, A. J. & Fürtbauer, I. (2017). Collective decision-making. In: International Encyclopedia of Primatology. Wiley-Liss. 

[53] King, A. J. & Fürtbauer, I. (2017). Social networks. In: International Encyclopedia of Primatology. Wiley-Liss.

[52] Fehlmann, G., O’Riain, M. J., Hopkins, P. W., O’Sullivan, J., Holton, M. D., Shepard, E. L. C., King, A. J. (2017) Identification of behaviours from accelerometer data in a wild social primate. Animal Biotelemetry 5:6. [R code] [Collar build guide] [Random forest workflow] [Trees and error rate] [Acceleration metrics] [BBC News] [Science Daily] [Cosmos] [IOL South Africa] [Discovery Channel Canada] 

[51] Couzin, I. D. & King, A. J. (2017) Group Movement, In: Reference Module in Life Sciences, Elsevier, ISBN: 978-0-12-809633-8.

[50] Fehlmann, G., O'Riain, J., Kerr-Smith, C.King, A. J. (2017) Adaptive space use by baboons (Papio ursnius) in response to management interventions in a human-changed landscape. Animal Conservation 20: 101–109. [Supplemental information/code]


[49] Duteil, M., Pope, E. C., Pérez-Escudero, A., de Polavieja, G. G., Fürtbauer, I., Brown, M. R., King, A. J. (2016) 

European sea bass show behavioural resilience to near-future ocean acidification. Royal Society Open Science 3(11), 160656 [Data] [Supplementary Material]

[48] Al Nahdi, A., Garcia De Leaniz, C., King, A. J. (2016) Spatio-temporal variation in length-weight relationships and condition of the ribbonfish Trichiurus lepturus (Linnaeus, 1758): Implications for fisheries management. PLoS ONE 11(8): e0161989. 

[47] Fehlmann, G.King, A. J. (2016) Bio-logging. Current Biology 26(18): pR830–R831.


[46] Al Nahdi, A., Garcia De Leaniz, C., King, A. J. (2016) Two distinct hyperostosis shapes in ribbonfish, Trichiurus lepturus (Linnaeus, 1758). Bulletin of the European Association of Fish Pathologists 36(3) 128-132.


[45] Hansen, M. J., Ward, A. J. W., Fürtbauer, I.*, King, A. J.* (2016) Environmental quality determines finder-joiner dynamics in socially foraging three-spined sticklebacks (Gasterosteus aculeatusBehavioral Ecology & Sociobiology 70: 889-899 *equal contribution



[44] King, A. J., Myatt, J. P., Fürtbauer, I., Oesch, N., Dunbar, R. I. M., Sumner, S., Usherwood, J. R., Brown, M. R. (2015) Social density processes regulate the functioning and performance of foraging human teams. Scientific Reports 5: 18260 [Video: summary]

[43] Johnson, C. J., Piel, A. K., Forman, D. W., Stewart, F. A., King, A. J. (2015) The ecological determinants of baboon troop movements at local and continental scales. Movement Ecology 3:14 [Additional file 1] [Additional file 2] [Additional file 3] [Additional file 4]

[42] Fürtbauer, I., King A. J., Heistermann, M. (2015) Visible Implant Elastomer (VIE) tagging and simulated predation risk elicit similar physiological stress responses in three-spined sticklebacks (Gasterosteus aculeatus). Journal of Fish Biology 86: 1644–1649  


[41] Fürtbauer, I., Pond, A., Heistermann, M., King A. J. (2015) Personality, plasticity, and predation: linking endocrine and behavioural reaction norms in stickleback fish. Functional Ecology 29: 931-940. [Video: example]

[40] King, A. J.*, Williams, L. J.*, & Mettke-Hofman, C. (2015) The effects of social conformity on Gouldian finch personality. Animal Behaviour 99: 25-31. *equal contribution  []


[39] Mamuneas, D., Spence, A. J., Manica, A., King, A. J. (2015) Bolder stickleback fish make faster decisions, but they are not less accurate. Behavioral Ecology 6: 91-96. [Data] [Video: example[BBC News Wales] [BEEP! podcast]



[38] Wilson, R. P., Richards, R., Hartnell, A., King, A. J., Piasecka, J., Gaihre, Y. K., Butt, T. (2014) A new approach to quantify semiochemical effects on insects based on energy landscapes. PLoS ONE 9(8): e106276   


[37] Strömbom. D., Mann, R. P., Wilson, A. M., Hailes, S., Morton, A. J., Sumpter, D. J. T., King, A. J. (2014) Solving the shepherding problem: Heuristics for herding autonomous, interacting agents. Journal of the Royal Society Interface (Top 10 most downloaded in 2014) 11: 20140719   [Movie 1] [Movie 2] [Movie 3[SUMMARY OF NEWS] e.g. [Time Magazine] [The Telegraph] [The Independent] [BBC News] [The Sydney Morning Herald] [Daily News] [Spiegel] Plus interviews on >15 Radio and TV shows



[36] King, A. J., Fürtbauer, I., Mamuneas, D.James, C., Manica, A. (2013). Sex-differences and temporal consistency in stickleback fish boldness. PLoS ONE 8(12): e81116  [Dataset] [Planet Earth] [Wales Online] [WIRED]


[35] Jolles, J. W., King, A. J., Manica, A., Thornton, A. (2013) Heterogeneous structure in mixed-species corvid flocks in flight. Animal Behaviour 85: 743–750.  [Planet Earth]

[34] Huchard, E., Charpentier, M. J. E., Marshall, H. H., King, A. J., Knapp, L. G., Cowlishaw, G. (2013) Paternal effects on access to resources in a promiscuous primate society. Behavioral Ecology 24: 229-236.  [Supplementary Information] [BBC Nature] [Planet Earth]


[33] Sueur, C., King, A. J., Pelé, M. Petit, O. (2013) Fast and accurate decisions as a result of scale-free network properties in two primate species. Springer Proceedings in Complexity 2013: 579-584. [New Scientist]

[32] King, A. J., Wilson, A. M., Wilshin, S. D., Lowe, J., Haddadi, H., Hailes, S., Morton, A. J. (2012). Selfish-herd behaviour of sheep under threat. Current Biology 22: R561-R562.  [Supplementary Model & Information] [Movie S1] [Movie S2] [Movie S3] widely reported by media, e.g. [Science] [BBC] [Scientific American] [Daily Telegraph] [Planet Earth] [ABC Science] [BBC Radio 4 - 23 mins]



[31] King, A. J., Cheng, L., Starke, S. D., Myatt, J. P. (2012) Is the true “wisdom of the crowd” to copy successful individuals? Biology Letters 8: 197-200.  [Sup.Figure.pdf] [] [Discover Magazine] [ScienceNews]


[30] Williams, L. J.King, A. J. & Mettke-Hofman, C. (2012) Colourful characters: Head colour reflects personality in a social bird, the Gouldian finch (Erythrura gouldiae). Animal Behaviour 84: 159-165.  [Planet Earth] [BBC Nature] [Daily Mail] [Cosmos Magazine] [Financial Times]


[29] Stevens, J. R. & King, A. J. (2012) The lives of others: Social rationality in animals. In: Simple heuristics in a social world. Edited by Ralph Hertwig, Ulrich Hoffrage, ABC Research Group. Oxford University Press


[28] Clark, F. E., Fitzpatrick, M., Hartley, A., King, A. J., Lee, T., Routh, A., Walker, S., & George, K. (2012). The relationship between behavior, adrenal activity and environment in zoo-housed Western lowland gorillas (Gorilla gorilla gorilla). Zoo Biology 31: 306-321. 


[27] Hobbs-Chell, H.King, A. J.Sharratt, H., Haddadi, H., Rudiger, S. R., Hailes, S., Morton, A. J., Wilson, A. M. (2012) Data-loggers carried on a harness do not adversely affect sheep locomotion. Research in Veterinary Science 93: 549–552. 


[26] King, A. J. & Sumpter, D. J. T. (2012) Murmurations. Current Biology 22: R112-R114. [The Huffington Post]


[25] Benavides, J., Huchard, E., Pettorelli, N., King, A. J., Brown, M. E., Archer, C. E., Appleton, C. C., Raymond, M., Cowlishaw, G. (2012) From parasite encounter to infection: multiple-scale drivers of parasite richness in a wild social primate population. American Journal of Physical Anthropology 147: 52-63. 



[24] King, A. J., Sueur, C., Huchard, E., Cowlishaw. G. (2011) A rule-of-thumb based on social affiliation explains collective movements in desert baboons. Animal Behaviour 82: 1337-1345.  [Basic model: NetLogo] [Planet Earth] [Earth Times] [EarthSky] [AskWhy!]


[23] Kaplan, B. S., O'Riain, J., van Eeden, R. & King A. J. (2011) A low-cost manipulation of food resources reduces spatial overlap between baboons (Papio ursinus) and humans in conflict. International Journal of Primatology 32: 1397-1412.  [Planet Earth]


[22] Pyritz, L. W., King A. J., Sueur, C. & Fichtel, C. F. (2011) Reaching a consensus: terminology and concepts used in coordination and decision-making research. International Journal of Primatology 32: 1268-1278. 


[21] King, A. J. & Sueur, C. (2011). Where next? Group coordination and collective decision-making by primates. International Journal of Primatology 32: 1245-1267. [Top cited paper in 2011-12


[20] Sueur, C. King, A. J., Conradt, L., Kerth, G., Lusseau, D., Mettke-Hoffmann, C., Schaffner, C.M., Williams, L. J., Zinner, D., Aureli, F. (2011) Collective decision-making and fission-fusion dynamics: a conceptual framework. Oikos 120: 1608-1617. 


[19] Sumner, S. & King, A. J. (2011). Actions speak louder than words in socially foraging human groups. Communicative & Integrative Biology 6: 755-757. 

[18] Haddadi, H., King, A. J., Wills, A. P., Fay, D., Lowe, J., Morton, A. J., Hailes, S. & Wilson, A. M. (2011) Determining association networks in social animals: choosing spatial-temporal criteria and sampling rates. Behavioral Ecology & Sociobiology 65: 1659-1668.  


[17] Sueur, C., Jacobs, A., Petit, O, Amblard, F. & King, A. J. (2011) How can social network analysis improve the study of primate behavior? American Journal of Primatology 73: 703-719. (Front cover).  

[16] King, A. J., Clark, F. E. & Cowlishaw, G. (2011) The dining etiquette of desert baboons: the roles of social bonds, kinship, and dominance in co-feeding networks. American Journal of Primatology 73: 768-774. [Planet Earth] [BBC Nature] 

[15] King, A. J.Narraway, C., Hodgson, L., Weatherill, A., Sommer, V. & Sumner, S. (2011) Performance of human groups in social foraging: the role of communication in consensus decision-making. Biology Letters 7: 237-240.  [The Economist]


[14] Katsikopoulos, K. & King, A. J. (2010) Swarm intelligence in animal groups: When can a collective out-perform an expert? PLoS ONE 5: e15505.   [Output of Model 1. xls] [Output of Model 3. xls]

[13] Couzin, I. D. & King, A. J. (2010) Animal group movement. In: Encyclopedia of Animal Behavior  Michael D. Breed and Janice Moore (Eds). Pages 25-32. Elsevier: Academic Press.


[12] King, A. J. (2010) Follow me! I'm a leader if you do; I'm a failed initiator if you don't? Behavioural Processes 84: 671-674.   


[11] King, A. J., Johnson, D. D. P. & Van Vugt, M. (2009) The origins and evolution of leadership. Current Biology 19: R911-R916.  [Science Daily]

[10] King, A. J., Isaac, N. J. B. & Cowlishaw, G. (2009) Ecological, social, and reproductive factors shape producer-scrounger dynamics in baboons. Behavioral Ecology 20: 1039-1049. (Cover image).  [Planet Earth]

[9] Huchard, E., Benavides, J., Setchell, J. M., Charpentier, M. J. E., Alvergne, A., King, A. J., Knapp, L. G., Cowlishaw, G., & Raymond, M. (2009) Studying shape in sexual signals: the case of primate sexual swellings. Behavioral Ecology & Sociobiology 63: 1231-1242.  


[8] King, A. J. & Cowlishaw, G. (2009c) All together now: behavioural synchrony in baboons. Animal Behaviour 78: 1381-1387.  [BBC Radio World Service] [BBC Radio 4 - Today Programme] [BBC Radio Wales]


[7] King, A. J. & Cowlishaw, G. (2009b) Leaders, followers and group decision-making. Communicative & Integrative Biology 2: 147-150. (Cover image).  

[6] King, A. J. & Cowlishaw, G. (2009a) Foraging opportunities drive interspecific associations between rock kestrels and desert baboons. Journal of Zoology 277: 111-118. (Cover image).  [Podcast]



[5] King, A. J. (2008) Leadership, coordinated behaviour, and information use in a social primate. PhD Thesis, UCL: London. [Abstract]


[4] King, A. J., Douglas, C. M. S., Isaac, N. J. B., Huchard, E. & Cowlishaw, G. (2008) Dominance and affiliation mediate despotism in a social primate. Current Biology 18: 1833-1838. (Cover image).  [Commentary] [Science Daily] [New Scientist]

[3] Clark, F. E. & King, A. J. (2008) A critical review of zoo-based olfactory enrichment. In: Chemical Signals in Vertebrates 11 Hurst, J. L., Beynon, R. J., Roberts, C. S. & Wyatt, T. D. (Eds), Springer.


[2] King, A. J. (2008) Dictators of the Desert. BBC Wildlife Magazine 26, 50-55. This article was also translated for Elethyrotipia magazine (Greece), and WWF Eko magazine (Sweden).


[1] King, A. J. & Cowlishaw, G. (2007) When to use social information: the advantage of large group size in individual decision-making. Biology Letters 3: 137–13.